in the more obtuse original definition:

(originally) Any unit of (originally cultural) information, such as a practice or idea, that is transmitted verbally or by repeated action from one mind to another in a comparable way to the transmission of genes.

as opposed to the more common definition you'd expect to find here...

(Internet) Media, usually humorous, which is copied and circulated online with slight adaptations, such as basic pictures, video templates, etc.

https://en.wiktionary.org/wiki/meme#Noun